| Pediatric Infectious Diseases Fellowship, Vanderbilt University, Nashville, TN | 1977 |
| Pediatric Infectious Diseases Fellowship, University of Washington, Seattle, WA | 1976 |
| Residency in Pediatrics, University of Washington, Seattle, WA | 1975 |
| Postdoctoral Fellowship, Division of Biology, California Institute of Technology, Pasadena, CA | 1964 |
| Postdoctoral Fellowship, Biochemistry Department, Columbia University, New York, NY | 1962 |
| Postdoctoral Fellowship, Division of Biology, California Institute of Technology, Pasadena, CA | 1961 |
| Pediatrics Internship, University of California Medical Center, San Francisco, CA | 1960 |
